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Kajihara, H., Grygier, M.J., Andrade, S.C., Bartolomaeus, T., Chernyshev, A.V., von Döhren, J., Gibson, R., Giribet, G., Hookabe, N., Junoy, J., Kvist, S., Maslakova, S.A., Mendes, C., Norenburg, J.L., Schwartz, M.L., Strand, M., Sun, S. & Turbeville, J.M. 2022. Case 3849 –Emplectonematidae Bürger, 1904 and Emplectonema Stimpson, 1857 (Nemertea, Monostilifera): proposed conservation of current usage by reversal of precedence of the family name with respect to Eunemertidae Joubin, 1894 and designation of a new type species for the genus. Bulletin of Zoological Nomenclature, 78: 184-200. PDF

Lynggaard, C., Oceguera-Figueroa, A., Kvist, S., Gilbert, M.T.P. & Bohmann, K. 2022. The potential of aquatic bloodfeeding and non-bloodfeeding leeches as a tool for iDNA characterisation. Molecular Ecology Resources, 22: 539-553. PDF


Darabi-Darestani, K., Sari, A., Khomenko, A., Kvist, S., Utevsky, S. 2021. DNA barcoding of Iranian leeches (Annelida: Clitellata: Hirudinida). Journal of Zoological Systematics and Evolutionary Research, 59: 1438-1452. PDF

Ceylan, M., Çetinkaya, O. & Kvist, S. 2021. The function of the waterfowl nests as reproduction and living areas for leeches (Annelida: Hirudinea). Animal Reproduction Science, 232: 106816. PDF

Iwama, R., Tessler, M., Siddall, M.E. & Kvist, S. 2021. The origin and evolution of antistasin-like proteins in leeches (Hirudinida, Clitellata). Genome Biology and Evolution, 13: evaa242. PDF


Anderson, K., Pepinelli, M., & Kvist, S. 2020. Natural occurrence of an experimental developmental phenotype in the leech Erpobdella punctata (Leidy, 1870) (Annelida: Clitellata: Hirudinea). Invertebrate Reproduction and Development, 64: 268-273. PDF

Williams, K.M., Barkdull, M., Fahmy, M., Hekkala, E., Siddall, M.E. &. Kvist, S. 2020. Caught red handed: iDNA points to wild source of CITES contraband leeches. European Journal of Wildlife Management, 66: 80. PDF

Sosa-Jiménez, V.M., Torres-Carrera, G., Manzano Marín, A., Kvist, S. & Oceuguera-Figueroa, A. 2020. Mitogenome of the blood feeding leech Haementeria acuecueyetzin (Hirudinida: Glossiphoniidae) from Tabasco, Mexico. Mitochondrial DNA Part B, 5: 3328-3330. PDF

Jiménez-Armenta, J., Kvist, S. & Oceguera-Figueroa, A. 2020. An exceptional case of mitochondrial tRNA duplication-deletion events in blood-feeding leeches. Organisms Diversity and Evolution, 20: 221-231. PDF

Kvist, S., Manzano-Marín, A., de Carle, D., Trontelj, P. & Siddall, M.E. 2020. Draft genome of the European medicinal leech Hirudo medicinalis (Annelida: Clitellata: Hirudiniformes) with emphasis on anticoagulants. Scientific Reports 10: 9885. PDF

Manglicmot, C., Oceguera-Figueroa, A., & Kvist, S. 2020. Bacterial endosymbionts of Placobdella (Annelida: Hirudinea: Glossiphoniidae): phylogeny, genetic distance, vertical transmission and co-speciation. Hydrobiologia, 847: 1177-1194. PDF

Anderson, K., Braoudakis, G., & Kvist, S. 2020. Genetic variation, pseudocryptic diversity, and phylogeny of Erpobdella (Annelida: Hirudinida: Erpobdelliformes), with emphasis on Canadian species. Molecular Phylogenetics and Evolution, 143: 106688. PDF


Iwama, R., de Carle, D., Manglicmot, C., Oceguera-Figueroa, A., Erséus, C., Siddall, M.E., & Kvist, S. 2019. Broad geographic sampling and DNA barcoding do not support the presence of Helobdella stagnalis (Linnaeus, 1758) in North America. Zootaxa 4671: 1-25. PDF

Mack, J., de Carle, D. & Kvist, S. 2019. Prey, populations, and the Pleistocene: evidence for low COI variation in a widespread North American leech. Mitochondrial DNA Part A, 30: 749-763. PDF

Phillips, A.J., Salas-Montiel, R., Kvist, S., & Oceguera-Figueroa, A. 2019. Phylogenetic position and description of a new species of “medicinal” leech from the eastern United States. Journal of Parasitology, 105: 587-597. PDF

Iwama, R., Oceguera-Figueroa, A., Giribet, G. & Kvist, S. 2019. The salivary transcriptome of Limnobdella mexicana (Annelida: Clitellata: Praobdellidae) and orthology determination of major leech anticoagulants. Parasitology, 146: 1338-1346. PDF

Mack, J. & Kvist, S. 2019. Improved geographic sampling provides further evidence for the separation of Glossiphonia complanata and Glossiphonia elegans (Annelida: Clitellata: Glossiphoniidae). Journal of Natural History, 53: 335-350. PDF

Khan, M. S. Guan, D.L., Kvist, S., Ma, L-B, Xie, J-Y., & Xu S-Q. 2019. Transcriptomics and differential gene expression in Whitmania pigra (Annelida: Clitellata: Hirudinida: Hirudinidae): contrasting feeding and fasting modes. Ecology & Evolution, 9: 4706-4719. PDF

Strand, M., Norenburg, J., Alfaya, J.E., Fernández-Álvarez, F.A., Andersson, H.S., Andrade, S.C.S., Bartolomeus, T., Beckers, P., Bigatti, G., Cherneva, I., Chernyshev, A., Chung, B.M., von Döhren, J., Giribet, G., Gonzalez-Cueto, J., Herrera-Bachiller, A., Hiebert, T., Hookabe, N., Junoy, J., Kajihara, H., Krämer, D., Kvist, S., Magarlamov, Y.T., Maslakova, S., Mendes, C.B., Okazaki, R., Sagorny, C., Schwartz, M., Sun, S-C., Sundberg, P., Turbeville, J.M., Xu, C-M. 2019. Nemertean taxonomy – implementing changes in the higher ranks, dismissing Anopla and Enopla. Zoologica Scripta, 48: 118-119. PDF


Kvist, S., de Carle, D., Cornejo, A. & Oceguera-Figueroa, A. Biological invasions or native ranges?: two curious cases of new distributional records in the Panama Canal. BioInvasions Records, in press. PDF

Koroiva, R. & Kvist, S. Estimating the barcoding gap in a global dataset of cox1 sequences for Odonata: close, but no cigar. Mitochondrial DNA, 29: 765-771. PDF

Tessler, M., Marancik, D., Champagne II, D., Dove, A., Camus, A., Siddall, M.E., & Kvist, S. 2018. Marine leech anticoagulant diversity and evolution. Journal of Parasitology, 104: 210-220. PDF

Kvist S. & Erséus, C. 2018. Two new European species of the marine genus Tubificoides (Annelida: Clitallata: Naididae) with notes on the morphology of T. pseudogaster (Dahl, 1960). Zootaxa, 4433: 561-573. PDF

Langer, S., Vezsenyi, K., de Carle, D., Beresford, D. & Kvist, S. 2018. Leeches (Annelida: Hirudinea) from the far north of Ontario: distribution, diversity, and diagnostics. Canadian Journal of Zoology, 96: 141-152. PDF


Koroiva, R., de Oliveira Roque, F., Pepinelli, M., Rodrigues, M. E., Lorenz-Lemke, A. P. & Kvist, S. 2017. DNA barcoding of odonates from the Upper Plata basin: database creation and genetic diversity estimation. PLoS ONE, 12: e0182283. PDF

de Carle D., Oceguera-Figueroa, A., Tessler, M., Siddall, M.E. & Kvist, S. 2017. Phylogenetic analysis of Placobdella (Hirudinea: Rhynchobdellida: Glossiphoniidae) with consideration of COI variation. Molecular Phylogenetics and Evolution, 114: 234-248. PDF

Kvist, S., Oceguera-Figueroa, A., Tessler, M., Jiménez-Armenta, J., Freeman Jr., R.M., Giribet, G. & Siddall, M.E. 2017. When predator becomes prey: investigating the salivary transcriptome of the shark-feeding leech Pontobdella macrothela (Hirudinea: Piscicolidae). Zoological Journal of the Linnean Society, 179: 725-737. PDF


Ceríaco, L.M.P. et al. (493 signatories). 2016. Photography-based taxonomy is inadequate, unnecessary, and potentially harmful for biological sciences. Zootaxa, 4196: 435-445. PDF

Pérez-Flores, J., Rueda-Calderon, H., Kvist, S., Siddall, M.E. & Oceguera-Figueroa, A. 2016. From the worm in the bottle of mezcal: iDNA confirmation of a leech parasitizing the Antillean Manatee. Journal of Parasitology, 102: 553-555. PDF

Wang, T., White, M.M., Kvist, S., & Moncalvo J-M. 2016. Genome-wide survey of gut fungi (Harpellales) reveals the first horizontally transferred ubiquitin gene from a mosquito host. Molecular Biology and Evolution, 33: 2544-2554. PDF

Sundberg, P., Kvist, S., & Strand, M. 2016. Evaluating the utility of single-locus DNA barcoding for the identification of ribbon worms (phylum Nemertea). PLoS ONE 11: e0155541. PDF

Sundberg, P., Andrade S.C., Bartolomeus, T., Beckers, P., von Döhren, J., Gibson, R., Giribet, G., Herrera-Bachiller, A., Junoy, J., Kajihara, H., Kvist, S., Kånneby, T., Krämer, D., Sun, S., Turbeville, J.M. & Strand, M. 2016. The future of nemertean taxonomy (phylum Nemertea) – a proposal. Zoologica Scripta, 45: 579-582. PDF

Oceguera-Figueroa, A., Manzano-Marin, A., Kvist, S., Moya, A., Siddall, M.E. & Latorre, A. 2016. Comparative mitogenomics of leeches (Annelida: Clitellata): genome conservation and Placobdella-specific trnD gene duplication. PLoS ONE 11: e0155441. PDF

Siddall, M.E., Brugler, M. & Kvist, S. 2016. Comparative transcriptomic analyses of 3 species of Placobdella (Rhynchobdellida: Glossiphoniidae) confirms a single origin of blood feeding in leeches. Journal of Parasitology 102: 143-150. PDF

Kvist, S. 2016. Does a global DNA barcoding gap exist in Annelida? Mitochondrial DNA 27: 2241-2252. PDF


Kvist, S., Chernyshev, A.V. & Giribet, G. 2015. Phylogeny of Nemertea with special interest in the placement of diversity from Far East Russia and northeast Asia. Hydrobiologia 760: 105-119. PDF

Herrera, A., Kvist, S., Giribet, G., & Junoy, J. 2015. Redescription of Micrura dellechiajei (Hubrecht, 1879) (Nemertea, Pilidiophora, Lineidae), a rare Mediterranean species. Journal of the Marine Biological Association of the UK 95: 1091-1100. PDF


Magesh, M., Kvist, S., & Glasby, C.J. 2014. Incipient speciation within the Namalycastis abiuma (Annelida: Nereididae) species group from southern India revealed by combined morphological and molecular data. Proceedings of the 11th International Polychaete Conference – Memoirs of the Museum Victoria 71: 169-176. PDF

Magesh, M., Glasby, C.J. & Kvist, S. 2014. Redescription of Namalycastsis glasbyi Fernando & Rajasekaran, 2007 (Annelida, Nereididae, Namanereidinae) from India. Proceedings of the Biological Society of Washington B 127: 455-465. PDF

Lenihan, J., Kvist, S., Fernández, R., Giribet, G. & Ziegler, A. 2014. A dataset comprising four micro-computed tomography scans of freshly fixed and museum earthworm specimens. GigaScience 3:6. PDF

Fernández, R., Kvist, S., Lenihan, J., Giribet, G. & Ziegler, A. 2014. Sine systemate chaos? A versatile tool for earthworm taxonomy: non-destructive imaging of freshly fixed and museum specimens using micro-computed tomography. PLoS One 9:e96617. PDF

Kvist, S., Laumer, C.L., Junoy, J. & Giribet, G. 2014. New insights into the phylogeny, systematics and DNA barcoding of Nemertea. Invertebrate Systematics 28: 287-308. PDF

Kvist, S., Brugler, M.R., Goh, T.G., Giribet, G. & Siddall, M.E. 2014. Pyrosequencing the salivary transcriptome of Haemadipsa interrupta (Annelida: Clitellata: Haemadipsidae): anticoagulant diversity and insight into the evolution of anticoagulation capabilities in leeches. Invertebrate Biology 133: 74-98. PDF


Kvist, S. 2013. Barcoding in the dark?: a critical view of the sufficiency of zoological DNA barcoding databases and a plea for broader integration of taxonomic knowledge. Molecular Phylogenetics and Evolution 69: 39-45. PDF

Kvist, S., Gi-Sik, M. & Siddall, M.E. 2013. Diversity and selective pressures of anticoagulants in three medicinal leeches (Hirudinida: Hirudinidae, Macrobdellidae). Ecology and Evolution 3: 918-933. PDF

Kvist, S. & Siddall, M.E. 2013. Phylogenomics of Annelida revisited: a cladistics approach using genome-wide EST data mining and examining the effects of missing data. Cladistics 29: 435-448. PDF


Magesh, M., Kvist, S. & Glasby, C.J. 2012. Description and phylogeny of Namalycastis jaya sp. n. (Polychaeta: Nereididae) from the southwest coast of India. Zookeys 238: 31-43. PDF

Siddall, M.E., Kvist, S., Phillips, A.J. & Oceguera-Figueroa, A. 2012. DNA barcoding of parasitic nematodes: is it kosher? Journal of Parasitology 98: 692-694.PDF


Kvist, S., Narechania, A., Oceguera-Figueroa, A., Fuks, B. & Siddall, M.E. 2011. Phylogenomics of Reichenowia parasitica, an alphaproteobacterial endosymbiont of the freshwater leech Placobdella parasitica. PLoS One 6: e28192. PDF

Kvist, S., Sarkar, I.N. & Siddall, M.E. 2011. Genome-wide search for leech antiplatelet proteins in the non-bloodfeeding leech Helobdella robusta (Rhyncobdellida: Glossiphoniidae) reveals evidence of secreted anticoagulants. Invertebrate Biology 130: 344-350. PDF

Kvist, S, Montanari, S.A., Yi, H., Fuks, B. & Siddall, M.E. 2011. Teaching biodiversity and evolutionary biology in a North American marine coastal environment. American Biology Teacher 73:72-77. PDF


Kvist, S., Oceguera-Figueroa, A., Siddall, M.E. & Erséus, C. 2010. Barcoding, types and the Hirudo-files: using information content to critically evaluate the identity of DNA barcodes. Mitochondrial DNA 21:198-205. PDF

Kvist, S., Sarkar, I.N. & Erséus, C. 2010. Genetic variation and phylogeny of the cosmopolitan marine genus Tubificoides (Annelida: Clitellata: Naididae: Tubificinae). Molecular Phylogenetics and Evolution 57:687-702. PDF

Oceguera-Figueroa, A., Kvist, S., Watson, S.C., Sankar, D., Overstreet, M. & Siddall, M.E. 2010. Leech collections from Washington State, USA, with the description of two new species of Placobdella (Annelida: Glossiphoniidae). American Museum Novitates No. 3701. PDF


Siddall, M.E., Fontanella, F.M., Watson, S.C., Kvist, S. & Erséus, C. 2009. Barcoding bamboozled by bacteria: convergence to metazoan mitochondrial primer targets by marine microbes. Systematic Biology 58: 445-451. PDF


Kvist, S., Dreyer, J. & Erséus, C. 2008. Two new species of Tubificoides (Annelida: Clitellata: Naididae) from the Blake Ridge methane seep in the North-west Atlantic Ocean. Proceedings of the Biological Society Washington B 121: 531-540. PDF


Erséus, C. & Kvist, S. 2007. COI-variation in Scandinavian marine species of Tubificoides (Annelida: Clitellata: Tubificidae). Journal of the Marine Biological Association of the UK 87: 1121-1126. PDF